Is Darius your vassal? I'm too tired to crunch the possible numbers, but the first thing that occurs to me is that it's actually his culture winning out in the outer ring, with the result that you get the tiles by virtue of being the master. That's not as satisfactory an explanation for the tiles toward Guimaraes as it is for those toward Persepolis, and less still for the inner ring itself, but maybe it's a step in the right direction.