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They are DDS texture files, you can find their locations in the civilization art defines XML. To edit them, you need a tool for DDS files. I think there are plugins for GIMP to do this, but the old school way is to use DXTBmp. Either way it gives you a normal image file you can edit however you want.

Note that editing the button DDS file does not change the flag shown on units in the game, you need to edit that one separately. It is a DDS file starting with FlagDECAL also referenced in the civilization art defines.
 
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